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  2. Ka/Ks ratio - Wikipedia

    en.wikipedia.org/wiki/Ka/Ks_ratio

    a. /K. s. ratio. In genetics, the Ka/Ks ratio, also known as ω or dN/dS ratio, [a] is used to estimate the balance between neutral mutations, purifying selection and beneficial mutations acting on a set of homologous protein-coding genes. It is calculated as the ratio of the number of nonsynonymous substitutions per non-synonymous site (K a ...

  3. 2-6-4 - Wikipedia

    en.wikipedia.org/wiki/2-6-4

    The first British examples of the 2-6-4T wheel arrangement were two locomotives built for the narrow-gauge Leek & Manifold Valley Light Railway in 1904. The first 4 ft in ( 1,435 mm) standard gauge examples were the class 1B of the Great Central Railway, built in 1914. [8] Richard Maunsell of the South Eastern and Chatham Railway (SECR ...

  4. LMS 3-Cylindered Stanier 2-6-4T - Wikipedia

    en.wikipedia.org/.../LMS_3-Cylindered_Stanier_2-6-4T

    The LMS Stanier Class 4P 3-Cylinder 2-6-4T is a class of steam locomotives designed for work over the London, Tilbury and Southend Railway route. All 37 were built in 1934 at Derby Works and were numbered 2500–2536. The third cylinder was provided to allow increased acceleration between the many stops on the L.T.&S.R. line.

  5. BR Standard Class 4 2-6-4T - Wikipedia

    en.wikipedia.org/wiki/BR_Standard_Class_4_2-6-4T

    BR Standard Class 4 2-6-4T engine no. 80118 at Guisborough railway station, 1950s. On the decision to build the BR standard series of locomotives, a series of class four tank engines was ordered, based on the ex- LMS Fairburn 2-6-4T with some modifications. The lineage of the class could therefore be tracked through the LMS/BR Class 4 2-6-4T ...

  6. 4-6-2+2-6-4 - Wikipedia

    en.wikipedia.org/wiki/4-6-2+2-6-4

    Evolved from. 2-6-2+2-6-2. Under the Whyte notation for the classification of steam locomotives by wheel arrangement, a 4-6-2+2-6-4 is a Garratt or Union Garratt articulated locomotive using a pair of 4-6-2 engine units back to back, with the boiler and cab suspended between them. The 4-6-2 wheel arrangement of each engine unit has four leading ...

  7. LMS 2-Cylindered Stanier 2-6-4T - Wikipedia

    en.wikipedia.org/.../LMS_2-Cylindered_Stanier_2-6-4T

    LMS: 4P. BR: 4MT. Withdrawn. 1960–1967. Disposition. All scrapped. Sir William Stanier 's London, Midland and Scottish Railway (LMS) Class 4P 2-Cylinder 2-6-4T was a class of 206 steam locomotive built between 1935 and 1943. They were based on his LMS 3-Cylinder 2-6-4T .

  8. 2-6-6-4 - Wikipedia

    en.wikipedia.org/wiki/2-6-6-4

    In the Whyte notation for the classification of steam locomotive wheel arrangement, a 2-6-6-4 is a locomotive with a two-wheel leading truck, two sets of six driving wheels, and a four-wheel trailing truck. All 2-6-6-4s are simple articulated locomotives . Other equivalent classifications are: UIC classification: (1'C)C2 ' (also known as German ...

  9. Type 4 Ka-Tsu - Wikipedia

    en.wikipedia.org/wiki/Type_4_Ka-Tsu

    Type 4 Ka-Tsu with machine guns mounted Rear-side angle view of Type 4 Ka-Tsu The Ka-Tsu's primary purpose was to transport cargo and/or troops ashore. [2] It had light armored shielding with a maximum of 10 mm. [1] Its engine compartment and electric final drives were hermetically sealed, as it was intended to be launched from a submarine. [3]

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